Gramma

San Diego Zoo’s Oldest Resident, Gramma the Tortoise, Dies at 141

Gramma, a beloved Galapagos tortoise and the oldest resident of the San Diego Zoo, has passed away at an estimated age of 141. Arriving at the zoo in either 1928 or 1931, Gramma charmed visitors with her gentle demeanor, living through numerous historical events. Her care specialists affectionately called her “the Queen of the Zoo” as she faced age-related health issues and was eventually euthanized. Galapagos tortoises are known for their longevity, with the oldest recorded, Harriet, living to 175 years old.

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